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The Permian Portal

Introduction

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Selected article on the Permian world and its legacies

Starfish or sea stars are echinoderms belonging to the class Asteroidea. About 1,500 living species of starfish occur on the seabed in all the world's oceans, from the tropics to subzero polar waters. They are found from the intertidal zone down to abyssal depths, 6,000 m (20,000 ft) below the surface.

Starfishes typically have a central disc and five arms, though some species have more than this. The aboral or upper surface may be smooth, granular or spiny, and is covered with overlapping plates. Starfish have tube feet operated by a hydraulic system and a mouth at the centre of the oral or lower surface. They are opportunistic feeders and are mostly predators on benthicinvertebrates. Several species having specialized feeding behaviours including eversion of their stomachs and suspension feeding. They have complex life cycles and can reproduce both sexually and asexually. Most can regenerate damaged parts or lost arms and they can shed arms as a means of defence. The Asteroidea occupy several significant ecological roles.

The fossil record for starfish is ancient, dating back to the Ordovician around 450 million years ago, but it is rather poor, as starfish tend to disintegrate after death. Only the ossicles and spines of the animal are likely to be preserved, making remains hard to locate. (see more...)

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Did you know?

... that the recently described synapsid Raranimus (pictured) is the most basal member of the order Therapsida, from which mammals are a descendant taxon?
... that the specific name of Tambachia trogallas, the type species of the trematopid temnospondyl Tambachia, refers to the Thuringian bratwurst that was frequently eaten by the describers of the species?
... that the Early Permian microsaur Rhynchonkos (pictured) shares many similarities with Eocaecilia, and may be an ancestor of caecilians?
... that the parareptile Eunotosaurus (restoration pictured) was once widely accepted as a transitional form between turtles and their prehistoric ancestors?
... that extinct diadectid reptiliomorphs (life restoration pictured) were the first large herbivorous land animals?

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Cross-bedded Permian Sandstone. These are the remnants of ancient desert sand dunes from around 300 to 350 million years ago.
Photo credit: Tony Atkin

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Selected article on the Permian in human science, culture and economics

Evolutionary thought, the conception that species change over time, has roots in antiquity. With the beginnings of biological taxonomy in the late 17th century, a new anti-Aristotelian approach to modern science challenged traditional essentialism. Naturalists began to focus on the variability of species; the emergence of paleontology with the concept of extinction further undermined the static view of nature. In the early 19th century, Jean-Baptiste Lamarck proposed the first fully formed theory of evolution.

In 1858, Charles Darwin and Alfred Russel Wallace published a new evolutionary theory that was explained in detail in Darwin's On the Origin of Species (1859). Unlike Lamarck, Darwin proposedcommon descent and a branching tree of life. The theory was based on the idea of natural selection, and it synthesized a broad range of evidence from animal husbandry, biogeography, geology, morphology, and embryology.

The debate over Darwin's work led to the rapid acceptance of the general concept of evolution, but the specific mechanism he proposed, natural selection, was not widely accepted until it was revived by developments in biology that occurred during the 1920s through the 1940s. Before that time most biologists argued that other factors were responsible for evolution. The synthesis of natural selection with Mendelian genetics during the 1920s and 1930s founded the new discipline of population genetics. Throughout the 1930s and 1940s, population genetics became integrated with other biological fields, resulting in a widely applicable theory of evolution that encompassed much of biology—the modern evolutionary synthesis. (see more...)

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